Love those pictures ........
'Lusitania' (the top one) was actually a Mark II, used at Arras in mid-1917. The original Mark I's were pretty tired out by then, so the Tank Corps had to use what they'd got. The Mark II's were superficially similar to the Mark I's (losing the tail wheels), but the key difference was that they were only designed for training, and used mild steel rather than boilerplate ! That's why the one at Bovington ('Flying Scotsman') is shot full of holes .........
I don't know anything about 'Escapade' (the second one), but the 'crew' are Germans, which could explain the grey / green tint
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And, as you said in your earlier post, this is a hornet's nest of a question. The splinter camo was definitely Mark I's only. The white / red / white recognition flashes were not introduced until mid-1918 (certainly not Cambrai), when (a) they needed to differentiate from the captured German Mark IV's, and (b) the newly-formed RAF were providing more ground support, so clear aerial recognition was important.
I think there is general acceptance that, post-1916, British tanks were basically brown. I'm not convinced there would have been a particular shade, as the Tank Corps was a very new formation, with a high attrition rate, and things just weren't that regimented. I suspect they used what was easily to hand, and let the mud do the rest. The Germans used captured stocks of French Railway paint on their tanks, and I'd imagine that the British would have done something similar / equally expedient.
So, in general, I don't think there's a definitive answer. That leaves everyone to do whatever they feel comfortable with (just don't do yellow with purple spots as my wife suggests !).
All the best,
Chris Hall
ADDENDUM 18/2/2015: I should have added that the usual word used to describe the shade of brown is 'chocolate'. It all depends, of course, how much cocoa and milk you like to use
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